Papers on social evolution & cooperation.

Krams, I., Kokko, H., Abolins-Abols, M., Krama, T. & Rantala, M.J. 2013. The excuse principle can maintain cooperation through forgivable defection in the prisoner’s dilemma game. Proceedings of the Royal Society of London B 280: 20131475.

Lehtonen, J. & Kokko, H. 2012. Positive feedback and alternative stable states in inbreeding, cooperation, sex roles and other evolutionary processes. Philosophical Transactions of the Royal Society of London B 367:211-221.

Fromhage, L. & Kokko, H. 2011. Monogamy and haplodiploidy act in synergy to promote the evolution of eusociality. Nature Communications 2:397.

Kokko, H. & Heubel, K.U. 2011. Prudent males, group adaptation, and the tragedy of the commons. Oikos 120: 641–656 (an invited Per Brinck Oikos Award article).

Jaatinen, K., Lehtonen, J. & Kokko, H. 2011. Strategy selection under conspecific brood parasitism: an integrative modelling approach. Behavioral Ecology 22:144-155.

Abbot et al. 2011. Inclusive fitness theory and eusociality. Nature 471:E1–E4.

López-Sepulcre, A., Norris, K. & Kokko, H. 2009. Reproductive conflict delays the recovery of an endangered social species. Journal of Animal Ecology 78:219-225.

Knopp, T., Heimovirta, M., Kokko, H. & Merilä, J. 2008. Do male moor frogs (Rana arvalis) lek with kin? Molecular Ecology 17:2522-2530.

Rankin, D.J., Bargum, K. & Kokko, H. 2007. The tragedy of the commons in evolutionary biology. Trends in Ecology & Evolution 22:643-651.

Kokko, H. 2007. Cooperative behaviour and cooperative breeding: what constitutes an explanation? Behavioural Processes 76:81-85.

Rankin, D.J., López-Sepulcre, A., Foster, K.R. & Kokko, H. 2007. Species-level selection reduces selfishness through competitive exclusion. Journal of Evolutionary Biology 20:1459-1468.

Sarhan, A. & Kokko, H. 2007. Multiple mating in the Glanville fritillary butterfly: a case of within-generation bet-hedging? Evolution 61:606-616.

Foster, K.R. & Kokko, H. 2006. Cheating can stabilise cooperation in mutualisms. Proceedings of the Royal Society of London B 273:2233-2239.

Kokko, H. 2003. Are reproductive skew models evolutionarily stable? Proceedings of the Royal Society of London B270:265-270.

Härdling, R. & Kokko, H. 2003. Life history traits as causes or consequences of social behaviour: why do cooperative breeders lay small clutches? Evolutionary Ecology Research 5:691-700.

Härdling, R., Kokko, H. & Arnold, K.E. 2003. Dynamics of the caring family. American Naturalist 161: 395-412.

Kokko, H. & Ekman, J. 2002. Delayed dispersal as a route to breeding: Territorial inheritance, 'safe havens' and ecological constraints. American Naturalist 160:468-484.

Kokko, H., Johnstone, R. A. & Wright, J. 2002. The evolution of parental and alloparental care in cooperatively breeding groups: when should helpers pay to stay? Behavioral Ecology 13:291-300.

López-Sepulcre, A. & Kokko, H. 2002. The role of kin recognition in the evolution of conspecific brood parasitism. Animal Behaviour 64:215-222.

Kokko, H., Johnstone, R.A. & Clutton-Brock, T.H. 2001. The evolution of cooperative breeding through group augmentation. Proceedings of the Royal Society of London B 268:187-196.

Kokko, H. & Lundberg, P. 2001. Dispersal, migration, and offspring retention in saturated habitats. American Naturalist 157:188-202.

Kokko, H. & Johnstone, R. A. 1999. Social queuing in animal societies: a dynamic model of reproductive skew. Proceedings of the Royal Society of London B 265:571-578.

Kokko, H. & Lindström, J. 1996. Kin selection and the evolution of leks: whose success do young males maximize? Proceedings of the Royal Society of London B 263:919-923.